On our coach we can see the tank where it's mounted in one bay next to the HydroHot. Coaches are different between models, but I'd think somewhere you should be able to see it. Our water tank is translucent so I can physically see how full it is, so you can verify things if you can hopefully find similar access, D.
If there's actually water in the tank, then it has to be a valve, or something floating in the tank that's blocked the intake, a highly unusual scenario. Check the proper positioning of any in-line shut offs like our coach has around the pump, but if you replaced the pump you likely already did that. The fact that the system was okay for awhile after filling is odd though, unless there's a blockage. A slow leak would do it, but your monitor and physical viewing would reveal that. A stuck open dump valve would be fairly obvious also; been there, done that because you have to remember to hold the dump switch for a second or two or it doesn't fully turn off or on.
As others have noted and discussed in previous threads here, there have been issues with the fill valve not switching correctly. If it didn't it would define your symptoms.
How about venting Guys? How would D check for a plugged tank vent? I know also it takes quite awhile at dewinterizing time to prime my pump to full flow, but given the entirety of his symptoms, priming is probably not the problem.
Joel